Perennial herbs or shrubs. Leaf blades orbicular, ovate or lanceolate, margins serrate or crenate. Flowers solitary in leaf axils. Epicalyx absent. Calyx deeply 5 lobed, accrescent, inflated in fruit, lobes apiculate, overlapping. Corolla yellow with or without a purple spot at the base. Staminal column included, filamentiferous at apex. Styles 8-14, stigmas capitate. Fruits schizocarpic. Mericarps 8-14, indehiscent, blackish, lateral walls reticulate-fenestrate, single celled, ventrally beaked, 1-seeded. Endoglossum absent.